Selection sorts what arrives. Covolution explains why some forms, variations, and trajectories are more likely to arrive than others.
Organisms and their molecular architectures structure the space in which variation occurs.
Protein folding funnels produce directional structural outcomes before selection acts on organismal fitness. (That does not abolish selection, but it means “design-like fit” can be partly prepared by physical and informational constraint.)
Natural selection is the best-established mechanism for cumulative adaptive fit, but the variation on which it acts is shaped by physical, developmental, and informational priors.
An informational architecture prior exists when outcome probabilities in a physical input-output map are significantly predicted by formal properties of the outputs, such as compressibility, modularity, degeneracy, or description length, beyond what is explained by immediate fitness differences alone.
In covolution theory, evolutionary change is decomposed into a selection term and a generative-architecture term. The selection term describes differential persistence of existing variants. The generative-architecture term describes biases in the production, accessibility, and stabilization of variants before selection. These biases may arise from genotype-phenotype map structure, developmental systems, mutation-rate architecture, epigenetic state, physical energy landscapes, symvironmental feedback, and predictive regulatory systems.
Covolution is a theory of structured arrival and recursive state-space construction, with natural selection retained as one filtering phase inside a larger information-processing loop.
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